Speciation

 The formation of a new species is called speciation.  A species is defined as a group of organisms that normally interbreed in nature to produce fertile offspring, and belong to the same gene pool.

An important part of forming a new species is separation of the gene pool of a species from the gene pool of the parent population.

Speciation can be separate into two types, based on the geographical relationship of the new population to its ancestral species:

  • Allopatric Speciation  (different areas)
  • Sympatric Speciation  (same habitat)

 

 Allopatric Speciation

It is when a species population is separated by a geographical barrier.  The species are known as allopatric populations.

This occurs in the following way:

  • Reduced selection pressure - a population moves into a new area free from competing species, predators, etc., with many ecological niches unfilled.
  • A population explosion follows which results in increased variation as most offspring survive, so alleles that were previously selected against are now free to be expressed.
  • Migration into new environments on the borders of the range gives rise to sub-species with different natural selection pressures.
  • Geographical barriers arise between sub-species and races, giving rise to geographical isolation.
  • The isolated populations will usually be exposed to different environmental selection pressures as they occupy different ranges.
  • Some of the isolated sub-species develop genetic and chromosomal differences that will no longer allow interbreeding with the parent population.  They are now genetically isolated and, thus, a new species.

 

 Allopatric speciation of squirrels in the Grand Canyon

The canyon is a barrier to dispersal by small mammals, and as a consequence the isolated populations can diverge.

 Sympatric Speciation

 Gene flow has been reduced between flies that feed on different food varieties, even though they both live in the same geographic area.